Check the transmission linkage to ensure that it is getting into gear. I would also check the transfer case linkage to make sure it isn't in neutral as well.
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When working with GEMS vehicles. Never try to locate a misfire by pulling plug leads off - pull the injector connector instead. Otherwise you can blow up the ECU.
Secondly never swap ECUs between vehicles to test them, because both vehicles and ECUs may stop working, and you'll have to have them all reset by a franchised LR dealer (ouch!).